The plot revolves around four +Anima children: Cooro, Husky, Senri, and Nana. Brought together by their +Anima powers, they search for others like themselves and a place to belong. Along the way, they encounter villains and friends alike, as well as other +Anima. Like many episodic adventure manga, there is not a big overarching story, rather the group travels from town to town, and at each stop over, they get wrapped up in local events. Examples of these adventures include Cooro helping a man fix and use a hang glider (man-lifting kite) to acquire medicine for his village, the group learning the secret of a monstrous +Anima that has been terrorizing another village, and Husky rescuing a man who has fallen in love with the legend of a mermaid. As the series progresses, more is learned about the characters' backstory, and about the setting of +Anima.
The world takes place on a fictional island continent, split between two nations: Astaria and Sailand. Both are mostly desert countries, but Astaria has more varied environments, with some steppes and forest. Between them is the great Moss Mountain range that keeps the two nations separate. On this range live the independent Kim-un-kur tribes. The majority of the series takes place in Astaria, with some later adventures taking place in the Moss Mountains and Sailand.
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